Paleontology

古生物学
  • 文章类型: Journal Article
    全球,在许多地质构造中都发现了硅化木。重要的是,木材的有机材料不再占主导地位;几乎所有的木化石都被矿化成无机二氧化硅材料。必须了解这些独特的地质过程,以更好地了解有机材料的石化,特别是在微米级。因此,我们的目的是使用全面的微观分析表征硅化木材的组成。使用的方法是XRF,ICP-MS,XRD,FTIR,和FE-EPMA。标本来自Jasinga,西爪哇,印度尼西亚。结果表明,木材硅化是通过二氧化硅从基质岩石渗入木材结构空间来控制的。在Jasinga,它们由上新世凝灰岩沉积岩控制。二氧化硅相的比例揭示了硅化程度的趋势。除了二氧化硅,微量元素的分布也表明了木化石与寄主岩石之间的地球化学相互作用。木化石受到通过硅化作用逐渐用二氧化硅代替有机碳基材料的影响。二氧化硅富集发生在木材内部,有利于矿化和重结晶。二氧化硅代替有机材料并保留木结构。微观分析方法提供了关于木材石化的全面观点,为古生物学研究带来更好的见解。
    Worldwide, silicified woods are found in many geological formations. Significantly, the organic materials of wood are no longer dominant; almost all wood fossils have been mineralized into inorganic silica materials. These unique geological processes must be understood to develop better understanding on organic material fossilization, particularly in the micron scale. Therefore, our aim was to characterize the composition of silicified wood using comprehensive microanalysis. The methods utilized were XRF, ICP-MS, XRD, FTIR, and FE-EPMA. Specimens are from Jasinga, West Java, Indonesia. The results showed that wood silicification was controlled by the infiltration of silica from the host rock into the spaces of the wood structure. In Jasinga, they are controlled by Pliocene tuffaceous sedimentary rocks. The ratio of silica phases revealed a trend in the degree of silicification. Besides silica, the distribution of trace elements also demonstrates the geochemical interaction between the wood fossil and host rock. Wood fossils are affected by the gradual replacement of organic carbon-based materials with silica through silicification. Silica enrichment occurs in the internal of wood, facilitates permineralization and recrystallization. Silica replaces organic material and preserves the wood structures. The microanalytical approach provides comprehensive perspectives on wood petrification, leads to better insights for paleontological studies.
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  • 文章类型: Journal Article
    自晚中生代以来,捕食驱动的中生代海洋革命(MMR)被认为已引起许多浅海无脊椎动物的测深分布发生了巨大变化。例如,在浅海环境中,有茎的海百合-等分线(Isocrinida)经历了惊人的下降,如今,它们仅限于深海环境(深度低于100m)。然而,这种转变的时机和同步性是一个争论的问题。MMR的延迟发作和/或转变为逆行,调用了南大洋古近纪期间的低捕食群落结构。特别是,来自南半球的最新数据表明,对深海环境的等分线的环境限制可能发生在南极洲和澳大利亚周围的始新世末期,后来在新西兰的早期。这里,我们报告了从波兰的中新世(北半球,CentralParatethys)。因此,全球,这是最年轻的浅海茎类海百合的记录。这一发现表明,一些残存的茎类海百合可能已经能够在上世纪中期生活在浅水环境中,并进一步证实,在全球范围内,对近海环境的深度限制并不同步。
    The predation-driven Mesozoic marine revolution (MMR) is believed to have induced a dramatic change in the bathymetric distribution of many shallow marine invertebrates since the late Mesozoic. For instance, stalked crinoids - isocrinids (Isocrinida) have undergone a striking decline in shallow-sea environments and today they are restricted to deep-sea settings (below 100 m depth). However, the timing and synchronicity of this shift are a matter of debate. A delayed onset of MMR and/or shifts to a retrograde, low-predation community structure during the Paleogene in the Southern Ocean were invoked. In particular, recent data from the Southern Hemisphere suggest that the environmental restriction of isocrinids to the deep-sea settings may have occurred at the end of the Eocene around Antarctica and Australia, and later in the early Miocene in New Zealand. Here, we report the anomalous occurrence of the isocrinids in shallow nearshore marine facies from the middle Miocene of Poland (Northern Hemisphere, Central Paratethys). Thus, globally, this is the youngest record of shallow-sea stalked crinoids. This finding suggests that some relict stalked crinoids may have been able to live in the shallow-water environments by the middle Miocene, and further confirms that the depth restriction of isocrinids to offshore environments was not synchronous on a global scale.
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  • 文章类型: Journal Article
    埃迪卡拉-寒武纪后生辐射的驱动因素尚不清楚,记录的保真度也是如此。我们使用全球年龄框架[5.8-5.1亿年前(Ma)]来估算海洋沉积岩体积和面积的变化,重建的生物多样性(平均属丰富度),和采样强度,与碳酸盐碳同位素(δ13Ccarb)和全球氧化还原数据[碳酸盐铀同位素(δ238Ucarb)]整合。采样强度与>535Ma前的总体平均重建生物多样性相关,而二阶(〜10-80Ma)全球海相-回归循环控制着不同海相沉积岩的分布。阿瓦隆组合的时间分布部分受深海硅质碎屑岩的时间和空间有限记录控制。定义阿瓦隆的后生形态群的每一次连续上升,白海,寒武纪组合似乎与δ13Ccarb最大值处的全球浅层海洋氧合事件相吻合,在重大海平面违规之前。虽然生物多样性的记录是有偏见的,早期的后生动物辐射和氧合事件与主要的海平面周期有关。
    The drivers of Ediacaran-Cambrian metazoan radiations remain unclear, as does the fidelity of the record. We use a global age framework [580-510 million years (Ma) ago] to estimate changes in marine sedimentary rock volume and area, reconstructed biodiversity (mean genus richness), and sampling intensity, integrated with carbonate carbon isotopes (δ13Ccarb) and global redox data [carbonate Uranium isotopes (δ238Ucarb)]. Sampling intensity correlates with overall mean reconstructed biodiversity >535 Ma ago, while second-order (~10-80 Ma) global transgressive-regressive cycles controlled the distribution of different marine sedimentary rocks. The temporal distribution of the Avalon assemblage is partly controlled by the temporally and spatially limited record of deep-marine siliciclastic rocks. Each successive rise of metazoan morphogroups that define the Avalon, White Sea, and Cambrian assemblages appears to coincide with global shallow marine oxygenation events at δ13Ccarb maxima, which precede major sea level transgressions. While the record of biodiversity is biased, early metazoan radiations and oxygenation events are linked to major sea level cycles.
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  • 文章类型: Journal Article
    南澳大利亚州的Emu湾页岩(EBS)在寒武纪Lagerstätten中异常,因为它捕获了在Burgess页岩型化石中罕见的解剖信息,由于其推断的近岸环境,其性质仍然存在争议。深入研究,将露头和钻孔数据与>25,000个化石标本的汇编相结合,揭示了EBS生物群在构造活跃的盆地内居住着扇三角洲复合体。在这种情况下保存软体生物是出乎意料的,这进一步突显了EBS与其他寒武纪Lagerstätten之间的差异。环境条件,包括氧气波动,边坡失稳,高悬浮泥沙浓度,和周期性的高能事件,除少数专业物种外,所有物种都抑制了下游前三角洲的定殖,但有利于下坡运输和其他主要特有物种的保护,浅水底栖动物.EBS提供了对冈比亚早期寒武纪动物多样性的非凡见解。这些结果证明了环境因素如何决定社区组成,并为理解这种独特的Konservat-Lagerstätte提供了框架。
    The Emu Bay Shale (EBS) of South Australia is anomalous among Cambrian Lagerstätten because it captures anatomical information that is rare in Burgess Shale-type fossils, and because of its inferred nearshore setting, the nature of which has remained controversial. Intensive study, combining outcrop and borehole data with a compilation of >25,000 fossil specimens, reveals that the EBS biota inhabited a fan delta complex within a tectonically active basin. Preservation of soft-bodied organisms in this setting is unexpected and further underscores differences between the EBS and other Cambrian Lagerstätten. Environmental conditions, including oxygen fluctuations, slope instability, high suspended sediment concentrations, and episodic high-energy events, inhibited colonization of the lower prodelta by all but a few specialist species but favored downslope transportation and preservation of other largely endemic, shallow-water benthos. The EBS provides extraordinary insight into early Cambrian animal diversity from Gondwana. These results demonstrate how environmental factors determined community composition and provide a framework for understanding this unique Konservat-Lagerstätte.
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  • 文章类型: Journal Article
    南美的最初人口是激烈的考古辩论的话题。最具争议的问题仍然是人类与大型动物互动的性质以及人类可能的作用,随着气候变化,在更新世末期几个大型哺乳动物属的灭绝中。在这项研究中,我们介绍了对化石遗骸的分析,这些化石遗骸属于新claerocalyptus(Xenarthra,Glyptodontidae),在Reconquista河岸发现的,Pampean地区(阿根廷)的东北部,其AMS14C测年对应于最后一次冰川最大值(21,090-20,811calYBP)。古环境重建,地层描述,骨材料的绝对年代,和沉积物表明,在雨季半干旱气候中,骨骼组合的埋葬事件相对较快。切割痕迹的定量和定性分析,屠宰序列的重建,对观察到的骨表面修饰中可能涉及的试剂的评估表明人类活动。我们的结果为讨论南美南部最早的人口提供了新的元素,特别是在最后一次冰川最大期间,人类与Pampean地区当地大型动物之间的相互作用。
    The initial peopling of South America is a topic of intense archaeological debate. Among the most contentious issues remain the nature of the human-megafauna interaction and the possible role of humans, along with climatic change, in the extinction of several megamammal genera at the end of the Pleistocene. In this study, we present the analysis of fossil remains with cutmarks belonging to a specimen of Neosclerocalyptus (Xenarthra, Glyptodontidae), found on the banks of the Reconquista River, northeast of the Pampean region (Argentina), whose AMS 14C dating corresponds to the Last Glacial Maximum (21,090-20,811 cal YBP). Paleoenvironmental reconstructions, stratigraphic descriptions, absolute chronological dating of bone materials, and deposits suggest a relatively rapid burial event of the bone assemblage in a semi-dry climate during a wet season. Quantitative and qualitative analyses of the cut marks, reconstruction of butchering sequences, and assessments of the possible agents involved in the observed bone surface modifications indicate anthropic activities. Our results provide new elements for discussing the earliest peopling of southern South America and specifically for the interaction between humans and local megafauna in the Pampean region during the Last Glacial Maximum.
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  • 文章类型: Journal Article
    高纬度峡湾可以作为沉积物陷阱,轴承不同类型的代理,从地球化学到微古生物学,使它们成为古环境重建的特殊工具。然而,可以存在一些非常规的代理,可以用来描述对过去变化的全面和详尽的解释。这里,研究埃迪斯托入口(罗斯海,南极洲)我们使用了不规则的棘突棘和蛇类(Ophionotusvictoriae)小骨来追踪最近3.6kyrsBP的环境变化。不规则的类孔虫可以作为有机物含量的代表,而O.维多利亚小骨可以用作稳定的海冰循环以及有机沉积事件的代表。O.维多利亚释放大量小骨,对人口的估计相当具有挑战性;尽管如此,存在数据,可以很容易地收集。通过将生成加法模型应用于这些数据的地层分布,我们检测到一个以前未被其他传统代理注意到的环境阶段:蛇形最佳(2-1.5kyrsBP)。总之,在这里,我们演示如何将棘皮动物的存在用作有价值的信息来源,同时证明了对二元数据进行建模以检测全新世地层记录中的长期趋势的潜力。
    High Latitude fjords can serve as sediment trap, bearing different type of proxies, from geochemical to micropaleontological ones, making them exceptional tools for paleoenvironmental reconstruction. However, some unconventional proxies can be present and can be used to depict a comprehensive and exhaustive interpretation of past changes. Here, studying a sediment core in Edisto Inlet (Ross Sea, Antarctica) we used irregular echinoid spines and ophiuroids (Ophionotus victoriae) ossicles to trace environmental changes throughout the last 3.6 kyrs BP. Irregular echinoids can serve as proxy for the organic matter content, while O. victoriae ossicles can be used as proxy for steady sea-ice cycle along with organic deposition events. O. victoriae release a high number of ossicles, making estimation about the population quite challenging; still, presence data, can be easily collected. By applying Generative Additive Models to the stratigraphical distribution of these data, we detected an environmental phase that was previously unnoticed by other traditional proxies: the Ophiuroid Optimum (2-1.5 kyrs BP). In conclusion, here we demonstrate how echinoderm presence can be used as a valuable source of information, while proving the potential of modelling binary data to detect long-term trend in Holocene stratigraphical records.
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  • 文章类型: Journal Article
    蛤虾是一群在三叠纪晚期繁荣昌盛的淡水甲壳类动物。在此期间,它们在整个盘古分布的陆相盆地的湖相沉积记录中很丰富。然而,它们显示出来自中部Pangea裂谷盆地和南部Gondwanan盆地的钳夹虾动物区系之间的显着分类学差异。在这一贡献中,我们展示了来自哥伦比亚东部科迪勒拉山脉(Bocas和Montebel地层)的湖相沉积演替的新的蛤虾化石组合,提供有关居住在西北冈瓦纳盆地的晚三叠世物种的信息。这项研究表明,冈瓦纳西北部的盆地与中央盘古的裂谷盆地共享Norian夹虾物种,并且它们的动物区系与冈瓦纳南部的盆地不同。此外,晚三叠纪蛤虾的古生物地理分布反映了该动物区系在整个Pangea中部裂谷中建立的河流-湖泊环境中的分布。因此,在中央Pangea早期破碎期间产生的裂谷可以充当分散的走廊。同时,裂谷还提供了古生物地理屏障,将中央Pangeaclam虾类动物与冈瓦纳南部隔离开来。
    Clam shrimps are a group of freshwater crustaceans who prospered during the Late Triassic. They were abundant in lacustrine sedimentary records of continental basins distributed throughout Pangea during this time. However, they show significant taxonomic differences between the clamp shrimp faunas from the rift basins of central Pangea and the southern Gondwanan basins. In this contribution, we show new fossil clam shrimp assemblages from the lacustrine sedimentary successions of the Eastern Cordillera of Colombia (the Bocas and Montebel formations), providing information on the Late Triassic species that inhabited the northwestern Gondwana basins. This study demonstrates that the basins of northwestern Gondwana shared Norian clamp shrimp species with rift basins of central Pangea and differed in their faunas with the basins of the southern portion of Gondwana. In addition, the Late Triassic clam shrimps paleobiogeographic distribution reflects the dispersal of this fauna throughout fluvial-lacustrine environments established in the rift valleys along the central Pangea. Therefore, the rift valleys produced during the early fragmentation of central Pangea could have acted as corridors for dispersion. Simultaneously, rift valleys also provided paleobiogeographic barriers that isolated the central Pangea clam shrimp faunas from southern Gondwana.
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  • 文章类型: Journal Article
    犹他州西南部圣乔治恐龙发现地点(SGDS)的下侏罗纪Moenave地层湖岸沉积物中的八个化石四足动物足迹无法分配给流行的恐龙(Anomoepus,Eubrontes,Gigandipus,施莱托,Kayentapus)或该地点的鳄鱼形(Batrachopus)ichnotaxa。三指和四指足迹不完整,由数字和数字尖端的印记组成。八个中的七个可能是pes指纹;其余的标本可能是Manus指纹。pes印刷品具有数字印记形态,并且与the的前投影和分叉角相似,一种主要存在于风尘古环境中的冰球,归因于新球藻突触。尽管它们的不完整性阻止了明确的转诊,尽管如此,SGDS曲目仍建议使用eucynodont曲目制造商,因此代表了一种罕见的曲目,在风成古环境之外,这种轨迹的早期中生代发生。
    Eight fossil tetrapod footprints from lake-shore deposits in the Lower Jurassic Moenave Formation at the St. George Dinosaur Discovery Site (SGDS) in southwestern Utah cannot be assigned to the prevalent dinosaurian (Anomoepus, Eubrontes, Gigandipus, Grallator, Kayentapus) or crocodyliform (Batrachopus) ichnotaxa at the site. The tridactyl and tetradactyl footprints are incomplete, consisting of digit- and digit-tip-only imprints. Seven of the eight are likely pes prints; the remaining specimen is a possible manus print. The pes prints have digit imprint morphologies and similar anterior projections and divarication angles to those of Brasilichnium, an ichnotaxon found primarily in eolian paleoenvironments attributed to eucynodont synapsids. Although their incompleteness prevents clear referral to Brasilichnium, the SGDS tracks nevertheless suggest a eucynodont track maker and thus represent a rare, Early Mesozoic occurrence of such tracks outside of an eolian paleoenvironment.
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  • 文章类型: Journal Article
    纽芬兰的Ediacaran保留了一些最古老的复杂宏观群落,其中一些以分形状的范围形态属Fractofusus为主。在这里,我们使用计算流体动力学和对Fractofususmisrai的详细重建来首次记录与这种沉积倾斜的生物相关的水动力现象及其与解释摄食策略相关的范围形态元素,解释最近记录的与电流倾斜的流变生长,并提供有关它们对Ediacaran海底影响的见解。斜向Fractofusus是常见的,可能代表最大化纵横比和最小化阻力之间的折衷。Fractofusus上表面的流动模式与溶解和细颗粒营养素的收集一致,以及气体交换。Fractofusus在下游产生尾流,证明倾斜的地形有可能通过允许细粒沉积物的沉积来改变古代海底的沉积模式。
    The Ediacaran of Newfoundland preserves some of the oldest complex macroscopic communities, several of which are dominated by the fractal-like rangeomorph genus Fractofusus. Here we use computational fluid dynamics and a detailed reconstruction of Fractofusus misrai to document for the first time hydrodynamic phenomena associated with this sediment-reclining organism and its rangeomorph elements that are relevant to interpreting feeding strategies, explain the recently documented rheotropic growth oblique to currents, and provide insights into their impact on the Ediacaran seafloor. Obliquely oriented Fractofusus are common, likely representing a compromise between maximized aspect ratio and minimization of drag. Flow patterns on the upper surface of Fractofusus are consistent with the collection of dissolved and finely particulate nutrients, as well as gas exchange. Fractofusus produce a wake downstream, demonstrating that reclining rangeomorphs had potential to modify sedimentation patterns on the ancient seafloor by potentially allowing deposition of fine-grained sediment.
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  • 文章类型: Journal Article
    埃迪卡拉-寒武纪过渡记录了动物多样化的关键阶段。全球化石记录记录了cloudinomorphs和其他带壳的管状生物的出现,其次是非生物矿化的小型碳质化石以及〜550至530Ma之间的高度多样化的小型架子化石。这里,我们报告了EdiacaranBocaina组的薄片和手样本中的各种微化石,巴西,分为五个描述性类别:细长实心结构(ES);细长填充结构(EF);两种类型的等维结构(EQ1和2)和带有盘绕末端的细长空心结构(CE)。这些标本,被解释为多样化的候选后生动物,早于上覆的Tamengo地层中Cloudina-Corumbella-Namacalathus生物区的最新Ediacaran生物矿化指数大化石。我们新的碳酸盐U-Pb年龄为Bocaina组,将这一新颖的化石记录定位在571±9Ma(加权平均年龄)。因此,我们的数据指向后生动物的多样化,包括生物矿化标本,让人联想到云纹切片,原牙,anabaritids,和hyolithids,除了动物的有机磷酸盐表面覆盖物,明显早于已知最早的骨骼化后生化石的记录。
    The Ediacaran-Cambrian transition documents a critical stage in the diversification of animals. The global fossil record documents the appearance of cloudinomorphs and other shelled tubular organisms followed by non-biomineralized small carbonaceous fossils and by the highly diversified small shelly fossils between ~ 550 and 530 Ma. Here, we report diverse microfossils in thin sections and hand samples from the Ediacaran Bocaina Formation, Brazil, separated into five descriptive categories: elongate solid structures (ES); elongate filled structures (EF); two types of equidimensional structures (EQ 1 and 2) and elongate hollow structures with coiled ends (CE). These specimens, interpreted as diversified candidate metazoans, predate the latest Ediacaran biomineralized index macrofossils of the Cloudina-Corumbella-Namacalathus biozone in the overlying Tamengo Formation. Our new carbonate U-Pb ages for the Bocaina Formation, position this novel fossil record at 571 ± 9 Ma (weighted mean age). Thus, our data point to diversification of metazoans, including biomineralized specimens reminiscent of sections of cloudinids, protoconodonts, anabaritids, and hyolithids, in addition to organo-phosphatic surficial coverings of animals, demonstrably earlier than the record of the earliest known skeletonized metazoan fossils.
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