paleontology

古生物学
  • 文章类型: Journal Article
    在这篇文章中,我们记录了在非鸟类恐龙的神经管中广泛存在的骨脊,包括种类繁多的蜥脚类动物,两个兽脚类动物,一个甲状腺素,还有一只鸭嘴龙.这些结构仅存在于尾椎中。它们是前后细长的,在运河的侧壁上发现,在分类学和序列上,大小和位置都不同。在现存的硬骨鱼中已经发现了类似的进入神经管的骨突起,Dipnoans,和urodelans,其中它们被认为是骨脊髓支撑。在大多数非哺乳动物中,围绕脊髓的硬脑膜与神经管的骨膜融合,支撑脊髓的齿状韧带可以穿过硬脑膜和骨膜直接固定在骨头上。这些结构在恐龙中的功能仍然不确定,但是在蜥脚类动物中,它们可能在尾巴的双侧运动和使用尾巴作为武器时稳定了脊髓。具有更广泛的意义,这项研究强调,通过仔细检查已知标本以前被忽视的特征,可以继续在总体解剖学水平上取得重要的新发现。
    In this article, we document the widespread presence of bony ridges in the neural canals of non-avian dinosaurs, including a wide diversity of sauropods, two theropods, a thyreophoran, and a hadrosaur. These structures are present only in the caudal vertebrae. They are anteroposteriorly elongate, found on the lateral walls of the canal, and vary in size and position both taxonomically and serially. Similar bony projections into the neural canal have been identified in extant teleosts, dipnoans, and urodelans, in which they are recognized as bony spinal cord supports. In most non-mammals, the dura mater that surrounds the spinal cord is fused to the periosteum of the neural canal, and the denticulate ligaments that support the spinal cord can pass through the dura and periosteum to anchor directly to bone. The function of these structures in dinosaurs remains uncertain, but in sauropods they might have stabilized the spinal cord during bilateral movement of the tail and use of the tail as a weapon. Of broader significance, this study emphasizes that important new discoveries at the gross anatomical level can continue to be made in part by closely examining previously overlooked features of known specimens.
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  • 文章类型: Journal Article
    背景:化石记录为观察数百万年的进化提供了独特的机会,但众所周知是不完整的。虽然不完整性在空间上有所不同,并且很难对经验部分进行估计,地质过程的计算机模拟可用于检查计算机中的不完整性的影响。我们结合了不同进化模式的模拟(停滞,(un)有偏差的随机游走)沉积碳酸盐岩平台地层,以检查从化石时间序列中恢复演化模式的程度,以及测试结果如何在碳酸盐岩台地的不同位置和由不同海平面曲线产生的多个地层结构之间变化。
    结果:沿陆上-海上梯度的地层结构和位置对通过统计检验恢复的演化模式只有很小的影响。对于随机游走的模拟,对正确模式的支持随着时间序列长度而减少。对谱系特征演变的视觉检查表明,而不是地层不完整,最大裂隙持续时间决定了化石时间序列与原始进化过程的差异。逐步定向演化更容易受到地层影响,把它变成断断续续的进化。相比之下,停滞仍然不受影响。
    结论:•化石时间序列有利于识别停滞和复杂,标点符号的进化模式。•不是地层不完整,但是稀有的存在,延长的缺口对性状进化的影响最大。这表明具有规则的中断频率和持续时间的不完整部分可以潜在地保留进化史而没有重大偏见。了解海平面波动等地层结构的外部控制对于区分地层效应和真正的演化过程至关重要。
    BACKGROUND: The fossil record provides the unique opportunity to observe evolution over millions of years, but is known to be incomplete. While incompleteness varies spatially and is hard to estimate for empirical sections, computer simulations of geological processes can be used to examine the effects of the incompleteness in silico. We combine simulations of different modes of evolution (stasis, (un)biased random walks) with deposition of carbonate platforms strata to examine how well the mode of evolution can be recovered from fossil time series, and how test results vary between different positions in the carbonate platform and multiple stratigraphic architectures generated by different sea level curves.
    RESULTS: Stratigraphic architecture and position along an onshore-offshore gradient has only a small influence on the mode of evolution recovered by statistical tests. For simulations of random walks, support for the correct mode decreases with time series length. Visual examination of trait evolution in lineages shows that rather than stratigraphic incompleteness, maximum hiatus duration determines how much fossil time series differ from the original evolutionary process. Gradual directional evolution is more susceptible to stratigraphic effects, turning it into punctuated evolution. In contrast, stasis remains unaffected.
    CONCLUSIONS: • Fossil time series favor the recognition of both stasis and complex, punctuated modes of evolution. • Not stratigraphic incompleteness, but the presence of rare, prolonged gaps has the largest effect on trait evolution. This suggests that incomplete sections with regular hiatus frequency and durations can potentially preserve evolutionary history without major biases. Understanding external controls on stratigraphic architectures such as sea level fluctuations is crucial for distinguishing between stratigraphic effects and genuine evolutionary process.
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  • 文章类型: Journal Article
    三叠纪是了解二叠纪末期物种大灭绝(EPME)后从古生代到中生代昆虫区系更替的关键时期;但是,早-中三叠世的化石矿床很少。在MonteSanGiorgio(瑞士)记录的保存完好的2.39亿年的昆虫化石动物群,这里展示了代表15个主要昆虫分支的248个化石。除了特殊的功能,包括它们的小尺寸和出色的保存,化石在该群体的进化史上具有重要意义。恢复了分类和生态多样性,包括淡水(蜻蜓和caddisfly)和陆生类群(真虫和黄蜂),证明了复杂的环境维持了一个由monurans主导的古社区(被认为没有在EPME中幸存下来),Midges,还有甲虫.有趣的是,还发现了一个带有外部卵皮的类似于布拉托的化石,对于理解古生代蟑螂向现存蟑螂的过渡和母体育龄的演变很重要。此外,发现了最年轻和第一个完整的Permithonidae标本和最古老的锯蝇化石。最后,圆形的身体,与精囊或番茄孢子相容,在几个蚊子样个体的腹部发现。如果这些是孢子,非种子植物可能是第一批昆虫植物,而不是裸子植物,就像最近所说的。总之,这些化石大大有助于理解昆虫的进化和古生代-中生代动物区系的周转。
    The Triassic represents a critical period for understanding the turnover of insect fauna from the Paleozoic to the Mesozoic following the end-Permian mass extinctions (EPME); however, fossil deposits from the Early-Middle Triassic are scarce. The exceptionally preserved 239 million-year-old fossil insect fauna recorded at Monte San Giorgio (Switzerland), including 248 fossils representing 15 major insect clades is presented here. Besides the exceptional features, including their small size and excellent preservation, the fossils have importance in the evolutionary history of the group. The taxonomic and ecological diversity recovered, including both freshwater (dragonflies and caddisflies) and terrestrial taxa (true bugs and wasps), demonstrates that complex environments sustained a paleocommunity dominated by monurans (thought not to have survived the EPME), midges, and beetles. Interestingly, a blattodean-like fossil bearing an external ootheca was also found, important for understanding Paleozoic roachoids to extant cockroaches\' transition and the evolution of maternal brood care. Moreover, the youngest and first complete specimen of †Permithonidae and the oldest sawfly fossils were discovered. Finally, round-shaped bodies, compatible with seminal capsules or lycophyte spores, were found on the abdomens of several midge-like individuals. If these are spores, non-seed-bearing plants could have been the first entomophilous plants rather than gymnosperms, as recently supposed. Altogether, these fossils contribute substantially to understanding insect evolution and Paleozoic-Mesozoic faunal turnover.
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  • 文章类型: Journal Article
    全球,在许多地质构造中都发现了硅化木。重要的是,木材的有机材料不再占主导地位;几乎所有的木化石都被矿化成无机二氧化硅材料。必须了解这些独特的地质过程,以更好地了解有机材料的石化,特别是在微米级。因此,我们的目的是使用全面的微观分析表征硅化木材的组成。使用的方法是XRF,ICP-MS,XRD,FTIR,和FE-EPMA。标本来自Jasinga,西爪哇,印度尼西亚。结果表明,木材硅化是通过二氧化硅从基质岩石渗入木材结构空间来控制的。在Jasinga,它们由上新世凝灰岩沉积岩控制。二氧化硅相的比例揭示了硅化程度的趋势。除了二氧化硅,微量元素的分布也表明了木化石与寄主岩石之间的地球化学相互作用。木化石受到通过硅化作用逐渐用二氧化硅代替有机碳基材料的影响。二氧化硅富集发生在木材内部,有利于矿化和重结晶。二氧化硅代替有机材料并保留木结构。微观分析方法提供了关于木材石化的全面观点,为古生物学研究带来更好的见解。
    Worldwide, silicified woods are found in many geological formations. Significantly, the organic materials of wood are no longer dominant; almost all wood fossils have been mineralized into inorganic silica materials. These unique geological processes must be understood to develop better understanding on organic material fossilization, particularly in the micron scale. Therefore, our aim was to characterize the composition of silicified wood using comprehensive microanalysis. The methods utilized were XRF, ICP-MS, XRD, FTIR, and FE-EPMA. Specimens are from Jasinga, West Java, Indonesia. The results showed that wood silicification was controlled by the infiltration of silica from the host rock into the spaces of the wood structure. In Jasinga, they are controlled by Pliocene tuffaceous sedimentary rocks. The ratio of silica phases revealed a trend in the degree of silicification. Besides silica, the distribution of trace elements also demonstrates the geochemical interaction between the wood fossil and host rock. Wood fossils are affected by the gradual replacement of organic carbon-based materials with silica through silicification. Silica enrichment occurs in the internal of wood, facilitates permineralization and recrystallization. Silica replaces organic material and preserves the wood structures. The microanalytical approach provides comprehensive perspectives on wood petrification, leads to better insights for paleontological studies.
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  • 文章类型: News
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  • 文章类型: Journal Article
    人工智能工具在taphonomy中是新的,并且正在快速增长。它们主要用于研究骨骼表面痕迹。为了调查这个问题,进行了文献计量学研究,以了解这个交叉领域的增长率,未来,和迄今为止的空白。从Scopus和谷歌学者元数据,用图表来描述数据,并利用普通最小二乘法进行回归统计。用词云进行探索性分析,主题建模,并使用论文的整个语料库进行了以潜在狄利克雷分配为方法的自然语言处理。从第一次注册到2023年,我们在Scopus上找到了8篇文章,在GoogleScholar上找到了32篇文章;大多数研究和引用最多的研究来自西班牙。从2016年到2018年,这些研究正在快速增长,回归表明未来几年可以保持增长。探索性分析表明,最常见的单词是标记,模型,数据,还有骨头.主题建模表明,研究高度集中在类似问题和解决这些问题的工具上,揭示了在分类学和古生物学中,还有更多的计算工具需要探索。
    Artificial intelligence tools are new in taphonomy and are growing fast. They are being used mainly to investigate bone surface marks. In order to investigate this subject, a bibliometric study was made to understand the growing rate of this intersectional field, the future, and gaps in the field until now. From Scopus and Google Scholar metadata, graphs were made to describe the data, and inferential statistics were made by regression with the Ordinary Least Squares method. Exploratory analysis with word clouds, topic modeling, and natural language processing with Latent Dirichlet Allocation as a method were also made using the entire corpus from the papers. From the first register until 2023, we found eight articles in Scopus and 32 in Google Scholar; the majority of the studies and the most cited were from Spain. The studies are growing fast from 2016 to 2018, and the regression shows that growth can be maintained in the coming years. Exploratory analysis shows the most frequent words are marks, models, data, and bone. Topic modeling shows that the studies are highly concentrated on similar problems and the tools to solve them, revealing that there is much more to explore with computational tools in taphonomy and paleontology as well.
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  • 文章类型: Journal Article
    Ediacaran生态系统的流体动力学模型说明了早期多细胞生物与环境水流之间的重要正反馈回路。因此,早期社区有助于化学塑造新的环境,在那里依赖氧气的生物可以茁壮成长。
    Fluid dynamics modeling of an Ediacaran ecosystem illustrates an important positive feedback loop between early multicellular organisms and environmental water flow. Early communities thus helped to chemically shape new environments where oxygen-dependent organisms could thrive.
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  • 文章类型: Journal Article
    自晚中生代以来,捕食驱动的中生代海洋革命(MMR)被认为已引起许多浅海无脊椎动物的测深分布发生了巨大变化。例如,在浅海环境中,有茎的海百合-等分线(Isocrinida)经历了惊人的下降,如今,它们仅限于深海环境(深度低于100m)。然而,这种转变的时机和同步性是一个争论的问题。MMR的延迟发作和/或转变为逆行,调用了南大洋古近纪期间的低捕食群落结构。特别是,来自南半球的最新数据表明,对深海环境的等分线的环境限制可能发生在南极洲和澳大利亚周围的始新世末期,后来在新西兰的早期。这里,我们报告了从波兰的中新世(北半球,CentralParatethys)。因此,全球,这是最年轻的浅海茎类海百合的记录。这一发现表明,一些残存的茎类海百合可能已经能够在上世纪中期生活在浅水环境中,并进一步证实,在全球范围内,对近海环境的深度限制并不同步。
    The predation-driven Mesozoic marine revolution (MMR) is believed to have induced a dramatic change in the bathymetric distribution of many shallow marine invertebrates since the late Mesozoic. For instance, stalked crinoids - isocrinids (Isocrinida) have undergone a striking decline in shallow-sea environments and today they are restricted to deep-sea settings (below 100 m depth). However, the timing and synchronicity of this shift are a matter of debate. A delayed onset of MMR and/or shifts to a retrograde, low-predation community structure during the Paleogene in the Southern Ocean were invoked. In particular, recent data from the Southern Hemisphere suggest that the environmental restriction of isocrinids to the deep-sea settings may have occurred at the end of the Eocene around Antarctica and Australia, and later in the early Miocene in New Zealand. Here, we report the anomalous occurrence of the isocrinids in shallow nearshore marine facies from the middle Miocene of Poland (Northern Hemisphere, Central Paratethys). Thus, globally, this is the youngest record of shallow-sea stalked crinoids. This finding suggests that some relict stalked crinoids may have been able to live in the shallow-water environments by the middle Miocene, and further confirms that the depth restriction of isocrinids to offshore environments was not synchronous on a global scale.
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  • 文章类型: Journal Article
    Most described Mesozoic ants belong to stem groups that existed only during the Cretaceous period. Previously, the earliest known crown ants were dated to the Turonian (Late Cretaceous, ca. 94-90 million years ago (Ma)) deposits found in the USA, Kazakhstan, and Botswana. However, the recent discovery of an alate male ant in Kachin amber from the earliest Cenomanian (ca. 99 Ma), representing a new genus and species, Antiquiformica alata, revises the narrative on ant diversification. Antiquiformica can be distinctly differentiated from all known male stem ants by its geniculate antennae with elongated scape, extending far beyond the occipital margin of the head and half the length of the funiculus, as well as its partly reduced forewing venation. Furthermore, the combination of a one-segmented waist with a well-developed node, elongated scape extending beyond the occipital margin, and reduced forewing venation, particularly the completely reduced m-cu and rs-m crossveins and absence of rm and mcu closed cells, firmly places the fossil within the extant subfamily Formicinae. Fourier transform infrared spectroscopy (FTIR) confirmed that the amber containing Antiquiformica alata originated from the Kachin mines in Myanmar. This discovery significantly revises our understanding of the early evolution of Formicinae. The presence of Antiquiformica in Cenomanian amber indicates that the subfamily Formicinae emerged at least by the start of the Late Cretaceous, with crown ants likely originating earlier during the earliest Cretaceous or possibly the Late Jurassic, although paleontological evidence is lacking to support the latter hypothesis.
    大多数被描述的中生代蚂蚁属于仅生存在白垩纪的基干类群。目前已知最古老的冠群蚂蚁发现于美国、哈萨克斯坦和博茨瓦纳的土仑阶沉积中(晚白垩纪,约94–90 Ma)。然而,在克钦琥珀中发现的一个森诺曼阶早期的有翅雄蚁新种(约99 Ma)——翼古老蚁( Antiquiformica alata)改变了对蚂蚁类群分化的认识。古老蚁属( Antiquiformica)与所有已知的雄性蚂蚁明显不同:触角呈膝状;柄节较长,延伸远超过后头边缘,长度约为鞭小节的一半;前翅翅脉部分减少。此外,古老蚁属( Antiquiformica)具有一个发育良好的腹结节、延伸超过后头边缘的细长柄节、简化的前翅翅脉,尤其是前翅横脉m-cu和rs-m完全消失、翅室rm和mcu不闭合,这些特征都表明翼古老蚁属于现存的蚁亚科。傅里叶红外光谱(FTIR)结果表明这件琥珀来自缅甸克钦矿区。新化石的发现极大地改变了我们对蚁亚科早期演化的认识。在克钦琥珀(森诺曼阶)中发现的古老蚁属( Antiquiformica)证实蚁亚科至少在晚白垩的早期就已经出现,而冠群蚂蚁出现时间肯定更早。冠群蚂蚁可能起源于白垩纪早期甚至侏罗纪晚期,尽管这一假设还未得到古生物学证据的支持。.
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  • 文章类型: Journal Article
    埃迪卡拉-寒武纪后生辐射的驱动因素尚不清楚,记录的保真度也是如此。我们使用全球年龄框架[5.8-5.1亿年前(Ma)]来估算海洋沉积岩体积和面积的变化,重建的生物多样性(平均属丰富度),和采样强度,与碳酸盐碳同位素(δ13Ccarb)和全球氧化还原数据[碳酸盐铀同位素(δ238Ucarb)]整合。采样强度与>535Ma前的总体平均重建生物多样性相关,而二阶(〜10-80Ma)全球海相-回归循环控制着不同海相沉积岩的分布。阿瓦隆组合的时间分布部分受深海硅质碎屑岩的时间和空间有限记录控制。定义阿瓦隆的后生形态群的每一次连续上升,白海,寒武纪组合似乎与δ13Ccarb最大值处的全球浅层海洋氧合事件相吻合,在重大海平面违规之前。虽然生物多样性的记录是有偏见的,早期的后生动物辐射和氧合事件与主要的海平面周期有关。
    The drivers of Ediacaran-Cambrian metazoan radiations remain unclear, as does the fidelity of the record. We use a global age framework [580-510 million years (Ma) ago] to estimate changes in marine sedimentary rock volume and area, reconstructed biodiversity (mean genus richness), and sampling intensity, integrated with carbonate carbon isotopes (δ13Ccarb) and global redox data [carbonate Uranium isotopes (δ238Ucarb)]. Sampling intensity correlates with overall mean reconstructed biodiversity >535 Ma ago, while second-order (~10-80 Ma) global transgressive-regressive cycles controlled the distribution of different marine sedimentary rocks. The temporal distribution of the Avalon assemblage is partly controlled by the temporally and spatially limited record of deep-marine siliciclastic rocks. Each successive rise of metazoan morphogroups that define the Avalon, White Sea, and Cambrian assemblages appears to coincide with global shallow marine oxygenation events at δ13Ccarb maxima, which precede major sea level transgressions. While the record of biodiversity is biased, early metazoan radiations and oxygenation events are linked to major sea level cycles.
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