Abstract：
We revise the Portuguese spiders of the genus Harpactea Bristowe, 1939 (Araneae: Dysderidae). The following seven new species are described: H. adicensis sp. nov., H. crespoi sp. nov., H. dolanskyi sp. nov., H. henriquesi sp. nov., H. korenkoi sp. nov., H. krejcii sp. nov. and H. pekari sp. nov. In three species, H. algarvensis Ferrández, 1990, H. minoccii Ferrández, 1982 and H. tavirensis Wunderlich, 2020 the females are described for the first time. The Portuguese Harpactea spiders belong to two species groups, namely the hombergi group and the corticalis group (sensu Deeleman-Reinhold 1993). The majority of the Portuguese representatives of the corticalis group probably constitute a monophyletic group endemic for the Iberian peninsula, the minoccii subgroup Ferrández, 1990. The synapomorphy of this clade is the unique position of partners during copulation: the male grasps one of the female\'s chelicerae between his chelicerae. The males\' chelicerae are morphologically adapted for this behaviour. Due to this unusual position during copulation the female genitalia are more distant from the male than in other Harpactea species. Therefore, the pedipalps are more elongated . Furthermore, the males of this subgroup possess larger basal part of male copulatory organ (the tegulum), where the ejaculate is stored before copulation. We observed that the males of this subgroup invest significantly more time into single copulation than the other representatives of the corticalis group with smaller tegulum. Therefore, enlargement of the tegulum might reflect different sperm competition strategy, in which males invest more ejaculate in each copulation. In Portugal, Harpactea spiders are frequently found under woody plants that produce slowly decomposing leaf litter, usually Quercus spp., or introduced Eucalyptus sp. These spiders require slightly humid substratum.
摘要：
我们修改了HarpacteaBristowe属的葡萄牙蜘蛛，1939年（Araneae：病菌科）。描述了以下七个新物种：H.adicensissp。11月。,H.crespoisp.11月。,H.dolanskyisp.11月。,H.henriquesisp.11月。,H.korenkoisp.11月。,H.krejciisp.11月。和H.pekarisp.11月。在三个物种中，H.algarvensisFrández,1990年，H.MinocciiFerrandez，1982年和H.tavirensisWunderlich，2020年女性首次被描述。葡萄牙热带蜘蛛属于两个物种，即hombergi组和皮层组(sensuDeeleman-Reinhold1993)。皮质皮层组的大多数葡萄牙代表可能构成伊比利亚半岛特有的单系组，Minoccii小组Ferrandez,1990.该进化枝的突触形态是交配过程中伴侣的独特位置：雄性在其chelicerae之间抓住雌性chelicerae之一。雄性龟科在形态上适应了这种行为。由于交配过程中的这种异常位置，雌性生殖器比其他Harpactea物种更远离雄性。因此,pedipalps更细长。此外，这个亚组的雄性具有较大的雄性交配器官的基底部分（tegulum），在交配前射精储存的地方。我们观察到，该亚组的雄性在单交配上投入的时间明显多于皮质小骨组的其他代表。因此,受精卵的扩大可能反映了不同的精子竞争策略，男性在每次交配中投入更多的射精。在葡萄牙,Harpactea蜘蛛经常在木本植物中发现，这些植物会产生缓慢分解的落叶，通常是蕨类植物。,或引入桉树。这些蜘蛛需要稍微潮湿的基质。
