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Abstract：
Diversity-biomass studies across (sub-)tropical forest strata have been explored, but our understanding on how multiple facets of forest diversity and abiotic factors regulate aboveground biomass across forest strata in temperate forests remains somehow unclear. Here, we conducted a comprehensive analysis of the effects and relative importance of taxonomic, phylogenetic and functional diversity, community-weighted mean (CWM) of trait values, coefficient of variation in individual tree diameter at breast height (CV DBH), and soil and topographic factors on aboveground biomass to select the diversity facets that could have the strongest response to abiotic factors across over- and understory and whole community in a temperate forest of South Korea. We used forest inventory, functional traits and environmental factors data from 259 plots to select the most important diversity facets with abiotic factors through multi-model inference tests, and we then used structural equation models. In the overstory, the most important regulators of aboveground biomass were topographic factor, species evenness, functional richness, and CV DBH. In the understory, the significant drivers of aboveground biomass were topographic factor, species evenness, phylogenetic species richness and CWM of height. In addition, diversity facets of overstory such as functional richness and CV DBH also had significant direct and/or indirect effects on understory aboveground biomass. Moreover, the diversity facets influencing aboveground biomass at the whole community were the combination of the multiple facets of forest diversity influencing aboveground biomass at each forest stratum. The role of functional dominance (CWM of height) seems to be negligible in the overstory but significant in the understory, indicating different diversity drivers as shown previously for a subtropical forest. Hence, our study suggests the urgent need of exploring diversity-biomass studies across forest strata in different forest ecosystems and types in order to provide more specific guidelines for the management of a specific natural forest.
摘要：
已经探索了跨（亚）热带森林地层的多样性-生物量研究，但是我们对森林多样性和非生物因素的多个方面如何调节温带森林中整个森林地层的地上生物量的理解仍然不清楚。这里,我们对分类学的影响和相对重要性进行了全面分析，系统发育和功能多样性，性状值的社区加权平均值(CWM)，个体树直径在胸高(CVDBH)时的变异系数，以及地上生物量上的土壤和地形因子，以选择可能对韩国温带森林的林上和林下以及整个社区的非生物因子反应最强的多样性方面。我们用了森林清查,来自259个地块的功能性状和环境因子数据，通过多模型推断测试，选择具有非生物因子的最重要的多样性方面，然后我们使用结构方程模型。在故事中，地上生物量最重要的调节剂是地形因子，物种均匀度，功能丰富，和CVDBH。在地下，地上生物量的重要驱动因素是地形因子，物种均匀度，系统发育物种丰富度和高度CWM。此外,林下的多样性方面，例如功能丰富度和CVDBH，也对林下地上生物量产生了显着的直接和/或间接影响。此外，影响整个群落地上生物量的多样性方面是影响每个森林地层地上生物量的森林多样性多个方面的组合。功能优势（身高的CWM）的作用在林下似乎可以忽略不计，但在林下却很重要。指示亚热带森林的不同多样性驱动因素。因此，我们的研究表明，迫切需要探索不同森林生态系统和类型的森林地层的多样性生物量研究，以便为特定天然林的管理提供更具体的指南。
